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Three thousand years ago, the Space Pirates Barban invaded Earth. The Starbeasts and the first Gingaman, warriors of the Ginga Forest, fought them with the mystical power called Earth and eventually were able to imprison them. The Ginga people later cloaked their forest within marked boundaries and passed on the duty of the Ginga warriors through generations. Hyuuga, Hayate, Gouki, Hikaru and Saya are chosen as the 133rd warriors of the Star Beast Swords. Ryouma is very happy that his elder brother Hyuuga succeeds to the honorable title. When Elder Ohghi holds the succession ceremony of Star Beast Swords, an earthquake breaks the seal of Barban. Ohghi orders the 133rd warriors to get the Ginga Braces hidden in Roaring Mountain. However, the Barban attacks them to prevent the birth of the new Gingamen. During the battle, Hyuuga got swallowed into a crack in the ground which Captain Zahab of the Barban created. Enraged, Ryouma activates his hidden Earth power and awaken the Ginga Braces. The Gingaman fights together with the Starbeasts against the Barban, who desire to revive the Demon Beast, Daitanix, on whose 'corpse' they built their castle.

Earth has regained peace but is attacked by a new enemy with space-time powers. Musashi, Cosmos' host tells the UPG that the Ultra Warriors along with his Ultra, Ultraman Cosmos, have been sealed inside an evil space-time castle by a warrior woman named Arina. The fate of the entire world will be at stake.

The series featured many firsts for the franchise, including the start of the story in a movie rather than the series proper. Ultraman Cosmos: The First Contact was the prequel to the series and took place ten years before episode 1. In it, 9-year-old Musashi Haruno encounters the being of light, Ultraman Cosmos, and befriends him as the two confront a threat to Earth. Ten years later, Musashi, now 19 and a member of Team EYES, once again encounters the being of light from his childhood. The dark being Chaos Header has appeared and is corrupting the monsters of Earth, making them become ravenous and violent creatures that threaten humanity. Through Musashi's will to see the beasts and humans live in peace, Cosmos grants him a new power that will allow him to heal the corrupted creatures.

Iron Claw is the leader of a global criminal empire known simply as "Crime". With a network of wealthy and influential sympathizers and employing an army of faceless, leather-masked thugs and cyborg assassins, Crime seeks to become the most powerful mafia organization in the world. To combat this threat, ISSIS, the International Science Special-Investigation Squad, is formed. ISSIS organizes its forces around the world to battle Crime, including Tokyo, Japan. Tokyo's ISSIS branch commander, Daisuke Kujirai, proposes a radical experiment to aid his forces in Japan. Taking the codename "Joker", he recruits four young test subjects to undergo his cyborg enhancement project. Surgically altered and given various energy manipulation powers and bionic enhancements, the four youths begin their mission to destroy Crime as the crime-busting quartet code-named J.A.K.Q. Dengekitai.